INDIANAPOLIS — Butler coach Chris Holtmann had the perfect plan to take down No. 1 Villanova on Wednesday night.

Play ugly, which worked beautifully.

Kethan Savage scored five of his 13 points during a decisive late run that helped the 18th-ranked Bulldogs end the nation’s longest winning streak at 20 games — and perhaps the Wildcats’ five-week run atop the polls — with a 66-58 victory that sent students streaming onto the court.

“We felt if we could make it ugly on the defensive end, we’d give ourselves a chance,” Holtmann said. “We did.”

What the Bulldogs (13-2, 2-1 Big East) also did was add another chapter to a storied program that captured the hearts of college basketball fans with back-to-back runs to the national championship game in 2010 and 2011 and broke hearts when two recent former players and the son of an assistant coach all died in 2016.

By extending their home-court winning streak to 12, the Bulldogs handed the defending national champs their first loss since last March. After losing 69-67 to Seton Hall in the Big East championship game, Villanova won six straight in the NCAA Tournament to capture its second national title. Then the Wildcats won their first 14 games this season for the longest winning streak in school history.

The loss left No. 2 Baylor , which beat Iowa State 65-63 on Wednesday night, and No. 5 Gonzaga as the nation’s only unbeaten teams.

“It stings right now,” Villanova preseason All-America Josh Hart said after scoring 13 points. “But we’re not going to be discouraged by it.”

Hart was 3 of 11 from the field and 1 of 5 on 3-pointers.

He wasn’t the only one having a rough night, though.

Kris Jenkins, the championship game hero, had 11 points. Villanova shot a season-low 37.3 percent from the field and was just 6 of 26 from beyond the arc. Coach Jay Wright picked up his first technical foul of the season.