CHENEY, Wash. — Hunter Wells fired a 6-yard touchdown pass to Kevin Rader in the end zone as time expired to lift Youngstown State to a 40-38 victory over Eastern Washington on Saturday to advance to the FCS national title game against James Madison.

Youngstown State (12-3) started the winning drive on its 42 with 4:24 left in the game, and used up the clock as it marched for the score.

Tevin McCaster rushed for 154 yards and three touchdowns for Youngstown State, which will be seeking its fifth national title.

Cooper Kupp caught 10 passes for 180 yards and two touchdowns for Eastern Washington (12-2), which saw its 11-game winning streak snapped.

The Penguins will play for the national championship Jan. 7 against James Madison, which beat North Dakota State 27-17 on Friday. The loss broke North Dakota State’s string of five consecutive FCS titles.

Saturday’s game was played in bitter cold, with the temperature dropping to single digits as the game progressed.

Youngstown State’s Jalyn Powell intercepted a Gage Gubrud pass on the first drive of the game. The Penguins marched down the field, with McCaster rushing in from the 4 for a 7-0 lead.

Eastern Washington replied with a drive that stalled on the 4, and settled for a 21-yard field goal by Roldan Alcobendas.

The Eagles scored again on Gubrud’s 4-yard touchdown pass to Kupp late in the first quarter for a 10-7 lead.

Gubrud completed 18 of 33 passes for 353 yards and three touchdowns, but suffered two interceptions.

Kupp is an All-America who won the Walter Payton Award last season as the top player in FCS and could become the second two-time winner when it is handed out Jan. 6.

Youngstown State running back Jody Webb ran for 101 yards, while Wells completed 16 of 24 passes for 244 yards and two touchdowns.